by JimDirt » Mon Apr 18, 2016 4:02 pm
What i am guessing you are experiencing is a stretched chain , when a chain stretches it does not ALL stretch , it depends on a lot of factors , like brand/quality of chain , and riding conditions , how its maintained , how its lubed all play a part in how and when and what part of a chain stretches
A brand new chain will start stretching from the minute you ride the bike , and it will wear the sprockets as it gets longer
(which is why your sprockets wear in a cup/hooked shape , because the longer the link gets the more space it needs to sit , since the sprocket is softer than the chain link , it wears the sprocket as the link's lengthen (actually its the rollers that elongate and wear not the sides of the link itself)
Now to your question..... If you happened to adjust the chain (prior to and after the rim swap) , and it happened to be in a tight/shorter part of the chain , or in a loose/stretched part of the chain , then it will affect how tight/loose it is at any given spot in rotation , so you could adjust it , and move the wheel a 1/4 turn , and it will be too tight or too loose now , even though it was fine when/where you adjusted it , if you did not check the entire rotation when you adjusted it , then it creates a issue with adjustment , on a stretched chain , you always adjust it at its tightest spot and hope the loosest spot is not so bad that the chain jumps the sprocket in the event of something hitting it , if it is noticeably different from tight to loose , then the chain (and usually sprockets , as they are a wear item and their longevity will be affected with a new chain , because they are "worn" to the old chain
No matter what , you will always have loose and tight spots in a chain , as torque is what does the stretching , along with improper adjusting or just lack of maintenance altogether **not implying this is the case with you** , my guess is the chain is either a cheaper chain , or its been on there a while (a season or more) , and this is the cause of the inaccuracy in tension
I swap wheels around on my 450R like you , for either practice and race wheels or trail riding , i buy $100+ chains (DID Gold Non O-Ring ER chains) about every 2 or 3 seasons , depending on how much i ride or how hard i ride each season , and i buy steel sprockets (because i am not worried about weight i am worried about strength and longevity)my chains wear very consistently , but when i notice the sprockets just barely cupping , i know its time and no adjustment is going to make it last any longer , its on a downhill slide just like once your valves start moving , its only a matter of time and your done
2020 CRF450R
2006 CRF450X

Weiser , Idaho